The California Highway Patrol said the shooting happened around 8:40 p.m. on the southbound 215 near Baseline Street.
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The driver reportedly pulled over and responding CHP officers found him inside the car with a gunshot wound.
The victim was loaded into an ambulance and transported to a hospital in unknown condition.
Footage from the scene showed a bullet hole in the windshield.
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An investigation is underway into whether the man was targeted or if it was a random shooting.